- Protein Name
- Dual specificity mitogen-activated protein kinase kinase 4
- Gene Name
- MAP2K4
- Organism
-
Homo sapiens
- Function
- Dual specificity protein kinase which acts as an essential component of the MAP kinase signal transduction pathway. Essential component of the stress-activated protein kinase/c-Jun N-terminal kinase (SAP/JNK) signaling pathway. With MAP2K7/MKK7, is the one of the only known kinase to directly activate the stress-activated protein kinase/c-Jun N-terminal kinases MAPK8/JNK1, MAPK9/JNK2 and MAPK10/JNK3. MAP2K4/MKK4 and MAP2K7/MKK7 both activate the JNKs by phosphorylation, but they differ in their preference for the phosphorylation site in the Thr-Pro-Tyr motif. MAP2K4 shows preference for phosphorylation of the Tyr residue and MAP2K7/MKK7 for the Thr residue. The phosphorylation of the Thr residue by MAP2K7/MKK7 seems to be the prerequisite for JNK activation at least in response to proinflammatory cytokines, while other stimuli activate both MAP2K4/MKK4 and MAP2K7/MKK7 which synergistically phosphorylate JNKs. MAP2K4 is required for maintaining peripheral lymphoid homeostasis. The MKK/JNK signaling pathway is also involved in mitochondrial death signaling pathway, including the release cytochrome c, leading to apoptosis. Whereas MAP2K7/MKK7 exclusively activates JNKs, MAP2K4/MKK4 additionally activates the p38 MAPKs MAPK11, MAPK12, MAPK13 and MAPK14.
Site |
Maximum Consensus Site Population |
Minimum Consensus Site Population |
Median Consensus Site Population |
MPP |
31 |
3 |
15.5 |
DRS |
19 |
3 |
12.0 |
MT3 |
16 |
9 |
12.5 |
AAS |
23 |
2 |
15.0 |
CMP |
22 |
3 |
10.0 |
PDIG |
2 |
2 |
2.0 |
ATP |
25 |
15 |
20.0 |
PDB ID |
Maximum Consensus Site Population |
MPP site |
DFG site |
PIF site |
DRS site |
DEF site |
EDI site |
MT3 site |
AAS site |
LBP site |
CMP site |
PDIG site |
ATP site |
PMP site |
Total Druggable Structures per Site |
2 |
0 |
0 |
1 |
0 |
0 |
1 |
3 |
0 |
2 |
0 |
1 |
0 |
3ALO
|
31 |
0 |
0 |
8 |
0 |
0 |
16 |
23 |
0 |
0 |
0 |
0 |
0 |
3ALN
|
25 |
0 |
0 |
19 |
0 |
0 |
9 |
18 |
0 |
22 |
2 |
25 |
0 |
3VUT
|
0 |
0 |
0 |
13 |
0 |
0 |
0 |
17 |
0 |
19 |
0 |
0 |
0 |